Although the threat of pests can be mitigated by the extensive use of pesticides, entire populations can be destroyed should a disease strike to which they have no resistance.
여기서 to which they have no resistance가 원래는 disease를 선행사로 하기 때문에 disease 뒤로 와야하는데 그냥 제일 뒤로 빠진 이유는 그냥 길어서 인거죠??